Review
Copyright ©2012 Baishideng Publishing Group Co., Limited. All rights reserved.
World J Orthop. Nov 18, 2012; 3(11): 190-198
Published online Nov 18, 2012. doi: 10.5312/wjo.v3.i11.190
Energy metabolism and the skeleton: Reciprocal interplay
Patrizia D Amelio, Anna Panico, Elena Spertino, Giovanni Carlo Isaia
Patrizia D Amelio, Anna Panico, Elena Spertino, Giovanni Carlo Isaia, Department of Surgical and Medical Disciplines, Gerontology Section, University of Torino, 10126 Torino, Italy
Author contributions: All authors contributed equally to this article.
Correspondence to: Elena Spertino, MD, Department of Surgical and Medical Disciplines, Gerontology Section, University of Torino, Cso Bramante 88, 10126 Torino, Italy. elenaspertino@gmail.com
Telephone: +39-11-6337140 Fax: +39-11-6961045
Received: March 14, 2011
Revised: July 31, 2012
Accepted: October 20, 2012
Published online: November 18, 2012
Abstract

The relation between bone remodelling and energy expenditure is an intriguing, and yet unexplained, challenge of the past ten years. In fact, it was only in the last few years that the skeleton was found to function, not only in its obvious roles of body support and protection, but also as an important part of the endocrine system. In particular, bone produces different hormones, like osteocalcin (OC), which influences energy expenditure in humans. The undercarboxylated form of OC has a reduced affinity for hydroxyapatite; hence it enters the systemic circulation more easily and exerts its metabolic functions for the proliferation of pancreatic β-cells, insulin secretion, sensitivity, and glucose tolerance. Leptin, a hormone synthesized by adipocytes, also has an effect on both bone remodelling and energy expenditure; in fact it inhibits appetite through hypothalamic influence and, in bone, stimulates osteoblastic differentiation and inhibits apoptosis. Leptin and serotonin exert opposite influences on bone mass accrual, but several features suggest that they might operate in the same pathway through a sympathetic tone. Serotonin, in fact, acts via two opposite pathways in controlling bone remodelling: central and peripheral. Serotonin product by the gastrointestinal tract (95%) augments bone formation by osteoblast, whereas brain-derived serotonin influences low bone mineral density and its decrease leads to an increase in bone resorption parameters. Finally, amylin (AMY) acts as a hormone that alters physiological responses related to feeding, and plays a role as a growth factor in bone. In vitro AMY stimulates the proliferation of osteoblasts, and osteoclast differentiation. Here we summarize the evidence that links energy expenditure and bone remodelling, with particular regard to humans.

Keywords: Leptin; Osteocalcin; Serotonin; Amylin; Bone mass; Energy metabolism