Published online Jan 27, 2022. doi: 10.4331/wjbc.v13.i1.15
Peer-review started: March 26, 2021
First decision: July 8, 2021
Revised: July 9, 2021
Accepted: January 13, 2022
Article in press: January 13, 2022
Published online: January 27, 2022
Processing time: 301 Days and 14.5 Hours
The genetically obese Zucker rat is hyperphagic and accumulates the excess of calorie intake in the form of lipids in spite of the insulin-resistant state of its adipocytes.
To investigate what biological events or natural biochemical processes drive the glucose utilization in fat cells of obese Zucker rats, which are not fully responsive to the lipogenic action of insulin.
Hydrogen peroxide is a biological chemical that can mimic several insulin actions on adipocytes, such as stimulating glucose entry and lipogenesis, and inhibiting lipolysis. Since it is a product of various enzymes in adipocytes, we focused our objective in searching whether the expression activity and biological effect of two types of them, namely, the monoamine oxidase (MAO) and semicarbazide sensitive amine oxidase (SSAO), abundant in adipocytes, were modified in obesity states.
Experimental methods included Zucker rat husbandry, with obese and lean littermates (the former bearing homozygotous recessive mutation of fa/fa gene), preparations of freshly isolated adipocytes, functional exploration of hexose transport using uptake assays with appropriate pharmacological agents, and determination of lipogenic activity, immunobloting, measurement of amine oxidase activities, and saturation binding analyses.
There is a good relationship between the increased binding capacity of tritiated-idazoxan and tritiated-(2-benzofuranyl)-2-imidazoline to imidazoline binding sites and the increased MAO-dependent tyramine oxidation in adipose tissue of obese rats. Stimulation of MAO or SSAO by their substrates in the presence of vanadate reproduced approximately two-thirds of the insulin stimulation of glucose uptake in fat cells. However, this insulin-like effect decreased as the insulin responsiveness of adipocytes decreased with obesity.
It cannot be stated whether the changes in MAO and SSAO expression are a cause or a consequence of the altered glucose handling in the fat cells of obese Zucker rats. At least, the increased tyramine oxidation is found in adipose tissues of the obese rats, not in the liver or in skeletal muscles, and can be associated with the dysregulation of the catecholaminergic system and of the energy balance found in that animal model of genetic obesity.
The increase of MAO activity in adipose tissue from obese rats has never been reported, at least to our knowledge. Among the limitations of our first description of an increased MAO activity in the adipose tissue of this animal model of obesity, is that we did not decipher the mechanisms supporting such up-regulation, and that it remains unclear whether elevated MAO is a cause or a consequence of the dysme