Original Article
Copyright ©2010 Baishideng Publishing Group Co., Limited. All rights reserved.
World J Gastroenterol. Oct 28, 2010; 16(40): 5047-5056
Published online Oct 28, 2010. doi: 10.3748/wjg.v16.i40.5047
Daily genetic profiling indicates JAK/STAT signaling promotes early hepatic stellate cell transdifferentiation
Ashley M Lakner, Cathy C Moore, Alyssa A Gulledge, Laura W Schrum
Ashley M Lakner, Cathy C Moore, Alyssa A Gulledge, Laura W Schrum, Department of Biology, University of North Carolina at Charlotte, Charlotte, NC 28223, United States
Laura W Schrum, Digestive and Metabolic Disorders Laboratory, Carolinas Medical Center, Charlotte, NC 28203, United States
Author contributions: Lakner AM and Moore CC performed the literature review, collected and analyzed the data, and drafted the manuscript, with major contributions from Schrum LW; Gulledge AA was responsible for generation of data and study design with guidance from Schrum LW; all authors read and approved the final manuscript.
Supported by National Institutes of Health Grant RO1 AA014891
Correspondence to: Laura W Schrum, PhD, Digestive and Metabolic Disorders Laboratory, Carolinas Medical Center, Charlotte, NC 28203, United States. laura.schrum@carolinashealthcare.org
Telephone: +1-704-3559670 Fax: +1-704-3557648
Received: June 10, 2010
Revised: June 30, 2010
Accepted: July 7, 2010
Published online: October 28, 2010
Abstract

AIM: To identify signaling pathways and genes that initiate and commit hepatic stellate cells (HSCs) to transdifferentiation.

METHODS: Primary HSCs were isolated from male Sprague-Dawley rats and cultured on plastic for 0-10 d. Gene expression was assessed daily (quiescent to day 10 culture-activation) by real time polymerase chain reaction and data clustered using AMADA software. The significance of JAK/STAT signaling to HSC transdifferentiation was determined by treating cells with a JAK2 inhibitor.

RESULTS: Genetic cluster analyses, based on expression of these 21 genes, showed similar expression profiles on days 1-3, days 5 and 6, and days 7-10, while freshly isolated cells (day Q) and day 4 cells were genotypically distinct from any of the other days. Additionally, gene expression clustering revealed strong upregulation of interleukin-6, JAK2 and STAT3 mRNA in the early stages of activation. Inhibition of the JAK/STAT signaling pathway impeded the morphological transdifferentiation of HSCs which correlated with decreased mRNA expression of several profibrotic genes including collagens, α-SMA, PDGFR and TGFβR.

CONCLUSION: These data demonstrate unique clustered genetic profiles during the daily progression of HSC transdifferentiation and that JAK/STAT signaling may be critical in the early stages of transdifferentiation.

Keywords: Cluster analysis; Fibrosis; Genetic profile; Hepatic stellate cell; Interleukin-6